England head coach Stuart Lancaster has cut five players from his World Cup squad ahead of a fortnight in a training camp in Denver, Colorado.

Maro Itoje, Elliot Daly, Ed Slater and Stephen Myler miss out, while Nick Easter has also been dropped as he continues to undergo treatment with club Harlequins on a back injury.

Aside from Easter, the players cut will be on a standby list as Lancaster, who will pick his final 31-man squad by the end of August, trims his 50-strong party to 45.

"I have spoken to all the players to explain that things can change very quickly and they can come back in at any point," said Lancaster.

"Nick will stay for treatment on his back rather than risk aggravating it further with travel and we look forward to him rejoining the squad in August when we will review the squad and also consider those who have been unable to begin the camp after suffering injuries at the end of last season.

"The players have worked very hard for the past three weeks and there is a really good feel about the group. Denver is the next stage in our training, a chance to develop further in a different environment."

England have spent three weeks at Penyhill Park.

England's 45-man training squad:

Chris Ashton (Saracens)
Dave Attwood (Bath Rugby)
Brad Barritt (Saracens)
Kieran Brookes (Northampton Saints)
Mike Brown (Harlequins)
Luther Burrell (Northampton Saints)
Sam Burgess (Bath Rugby)
Danny Care (Harlequins)
Danny Cipriani (Sale Sharks)
Calum Clark (Northampton Saints)
Dan Cole (Leicester Tigers)
Alex Corbisiero (Northampton Saints)
Luke Cowan-Dickie (Exeter Chiefs)
Lee Dickson (Northampton Saints)
Kyle Eastmond (Bath Rugby)
Owen Farrell (Saracens)
George Ford (Bath Rugby)
Alex Goode (Saracens)
Jamie George (Saracens)
James Haskell (Wasps)
Jonathan Joseph (Bath Rugby)
Courtney Lawes (Northampton Saints)
Joe Launchbury (Wasps)
George Kruis (Saracens)
Matt Kvesic (Gloucester Rugby)
Joe Marler (Harlequins)
Jonny May (Gloucester Rugby)
Ben Morgan (Gloucester Rugby)
Matt Mullan (Wasps)
Jack Nowell (Exeter Chiefs)
Geoff Parling (Exeter Chiefs)
Chris Robshaw (Harlequins)
Semesa Rokoduguni (Bath Rugby)
Henry Slade (Exeter Chiefs)
Billy Twelvetrees (Gloucester Rugby)
Mako Vunipola (Saracens)
Billy Vunipola (Saracens)
Anthony Watson (Bath Rugby)
Rob Webber (Bath Rugby)
Richard Wigglesworth (Saracens)
David Wilson (Bath Rugby)
Tom Wood (Northampton Saints)
Marland Yarde (Harlequins)
Ben Youngs (Leicester Tigers)
Tom Youngs (Leicester Tigers)